When two notorious scientists, working for The Light, arrived in Bwunda to conduct experiments on the local gorilla population, he was the first to be captured - by the League of Shadows - and experimented upon.
The scientists subjected Tolifhar to inhumane experiments, including the removal of his brain and its replacement with one of their own. This process resulted in the deaths of both the ape and the body of Delores Winters - the aged female scientist, who was as much a victim as Tolifhar himself, having had her own brain removed and her body hijacked by the mad scientist, The Ultra-Humanite years ago.
Surviving the experiment, Ultra-Humaite would go on to commit atrocities, both against humanity and the apes that Tolifhar had once held dear.
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